Most arthropods have a well-developed nervous system, containing a pair of ganglia and two long nerves down the esophagus, connecting the brain to a nerve cord that runs along the ventral part of the body. The ganglia serve as the main communication centers which coordinate the movement of the legs and the wings. There are several ganglia for each major body part.
Arthropods also contain simple sense organs such as statocysts and chemical receptors. Compound eyes are one of their main sensory organs, holding a myriad of separate lenses which can detect colour and movement extremely well, as well as ultraviolet light, which is invisible to humans.
Both crustaceans and insects have a well-developed sense of taste, smell, movement, and hearing. Taste receptors are located on their mouthparts, antennae and legs, while sensory hairs are situated all across the body. Insect ears are often found in unusual places, such as the eardrums of a grasshopper, which are located behind the legs.